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In the year 2022, the gaming industry has reached a new height after the development of the NerveGear, a helmet-like controller that allows the user to fully dive their senses into the world of virtual reality. After a closed beta with only 1,000 testers, the Virtual Reality MMORPG Sword Art Online is launched.

Gurren Lagann takes place in a fictional future where Earth is ruled by the Spiral King, Lordgenome, who forces mankind to live in isolated subterranean villages. These villages have no contact with the surface world or other villages, and are under constant threat of earthquakes.

Alucard is not just any immortal, but probably one of the oldest and strongest beings known to exist. This vampire, however, is duty bound to the Hellsing family, who acts as the dogs of England's queen, protecting the country from the most dangerous parts of the underground, namely other vampires.