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A young sorceress, Lucy Heartfilia, travels to the land of Fiore to join the magical Fairy Tail Guild. Along the way, she meets Natsu Dragneel, a teenage boy looking for a dragon named Igneel.

By its eleventh expansion pack, the massively multiplayer online role-playing game (MMORPG) Elder Tale has become a global success, having a following of millions of players. However, during the release of its twelfth expansion pack: Novasphere Pioneers, thirty thousand Japanese gamers who are all logged on at the time of the update, suddenly find themselves transported inside the game world and donning their in-game avatars.

Komatsu Nana moves to Tokyo, following after her boyfriend Shouji to gain a life she has always dreamed of. The anime is about two girls named Nana, who are very different.